Link to Sam's Google Scholar Page

*indicates undergraduate coauthor. = indcates authors contributed equally to work.


Tye SP, Fey SB, Gibert JP, Siepielski AM. 2024. Predator mass mortality events restructure freshwater food webs via trophic decoupling. Nature. 626: 335-340. Link  *Cover Image*  Research Briefing  Nature Podcast


Theus ME*, Michaels J, Fey SB. 2023. Interactive effects of temperature and Bisphenol A on phytoplankton growth and community structure. Conservation Physiology. 11: coad02. Link

Slein MA*, Bernhardt JR, O’Connor MI, Fey SB. 2023. Effects of thermal fluctuations on biological processes: a meta-analysis of experiments manipulating thermal variability. Proceedings of the Royal Society B: Biological Sciences. 290: 20222225. Link


Meier HS*, Schuman IJ*, Layden TJ, Ritz A, Kremer CT, Fey SB. Temperature mediated transgenerational plasticity influences movement behavior in the green algae Chlamydomonas reinhardtii. Functional Ecology. 36: 2969-2982. Link  *Cover Image*

Gerhard M, Koussoroplis AM, and 18 others including Fey SB. 2022. Environmental variability in aquatic ecosystems: avenues for future experimental studies in a multifactorial world. Limnology and Oceanography Letters. Early view Link

Tye SP, Siepielski AM, Bray A, Ryple AL, Phelps NBD, Fey SB. 2022. Climate warming amplifies frequency of fish mass mortality events across north temperate lakes. Limnology and Oceanography Letters.  7: 510-519. Link     Media Coverage: Science Friday

Schuman IJ*, Meier HS*, Layden TJ, Fey SB. 2022. The relationship between thermal spatial variability and mean temperature alters movement and population dynamics. Ecosphere. 13: e4254. Link

Theus ME*, Layden TJ, McWilliams N*, Crafton-Temple S*, Kremer CT, Fey SB. 2022. Photoperiod regimes influence the shape and scaling of freshwater phytoplankton responses to light and temperature. Oikos. e08839. Link

Shoemaker LG, Hallet LM, and 17 others including Fey SB. 2022.The long and the short of it: Decomposing synchrony and compensation across temporal scales. Ecology. 103: e3650. Link 


Fey SB=, Kremer CT=, Layden TJ, Vasseur DA. 2021. Resolving the consequences of gradual phenotypic plasticity for populations in variable environments. Ecological Monographs. 91: e01478. Link

Walter JA, Shoemaker LG, Lany NK, Castorani MCN, Fey SB, Dudney JC, Gherardi L, Reyes CP, Rypel AL, Cottingham KL, Suding KN, Reuman DC, Hallett LM. 2021. The spatial synchrony of species richness and its implications for ecosystem stability. Ecology.102: e03486. Link

Layden TJ, Kremer CT, Pietsch DB*, Kolk M*, Vasseur DA, Fey SB. 2021. Thermal acclimation influences the growth and toxin production of freshwater cyanobacteria. Limnology and Oceanography Letters. 7: 34-42. Link.


Fey SB, Theus ME*, Ramirez AR. 2020. Course-based undergraduate research experiences in a remote setting: Two case studies documenting implementation and student perceptions. Ecology and Evolution. 10: 12528-12541. Link.


Fey SB=, Gibert JP=, Siepielski AM. 2019. The consequences of mass mortality events for the structure and dynamics of biological communities. Oikos. 128, 1679-1690. Link. **Selected as ‘Editors’ Choice’ for October Issue**

AL Till*, AL Rypel, A Bray, SB Fey. Temperature extremes are coincident with fish die-offs in north temperate lakes. Nature Climate Change. 9, 637-641. Link   Media coverage: Reed MagazineWisconsin Public Radio, USA Today, Milwaukee Journal Sentinel, Frontiers in Ecology and the Environment

Fey SB=, Vasseur DA=, Alujevic K, Kroeker KJ, Logan ML, DeLong JP, O’Connor MI, Peacor S, Rudolf VHW,  Selden RL, Sih A, Clusella-Trullas S. Opportunities for Behavioral Rescue Under Rapid Environmental Change. Global Change Biology. 25, 3110-3120. Link   Media coverage: Frontiers in Ecology and the Environment


Woods HA, Kingsolver JG, Fey SB, Vasseur DA. Uncertainty in geographic estimates of performance and fitness. Methods in Ecology and Evolution. 9: 1996-2008Link

Culler LE, Wood ZT*, Diaz J, Fey SB, Ayres MP, Timmins D. Streams in an uninhabited watershed have predictably different thermal sensitivities to variable summer air temperatures. Freshwater Biology. 63: 676-686. Link

Kremer KT=, Fey SB=, Arellano AA*, Vasseur DA. Gradual plasticity alters population dynamics in variable environments: thermal acclimation in the green alga Chlamydomonas reinhartdii.  Proceedings of the Royal Society B: Biological Sciences. 285: 20171942. Link

Collins SL, Avolio ML, Gries C, Hallett LM, Koerner SE, La Pierre KJ, Rypel AL, Sokol ER, Fey SB, Flynn DFB, Jones SK, Ladwig LM, Ripplinger J, Jones MB. Temporal heterogeneity increases with spatial heterogeneity in ecological communities. Ecology. 99: 858-865. Link


Fey SB, Wieczynski D. 2017. The temporal structure of the environment may influence range expansions during climate warming. Global Change Biology23, 636-645. Link

Cottingham KL, Fey SB=, Fritschie KJ=, Trout-Haney JV=. 2017. Advancing ecosystem science by promoting greater use of theory and multiple research approaches in graduate education. Ecosystems. 20, 267-273. Link 


Dillon ME, Woods A, Wang G, Fey SB, Vasseur DA, Telemeco RS, Marshall KE, Pincebourde S. 2016. Life in the frequency domain: the biological impacts of changes in climate variability at multiple time scales. Integrative and Comparative Biology. 56, 14-30. Link

Fey SB, Vasseur DA. 2016. Thermal variability alters the impact of climate warming on consumer-resource systems. Ecology. 97, 1690-1699. Link


Sanders RW, Cooke SL, Fischer JM, Fey SB, Heinze AW, Jeffrey WH, Macaluso AL, Moeller RE, Morris DP, Neale P, Olson M, Pakulsk JD, Porter JA, Schoener DM, Williamson CE. 2015. Shifts in plankton community structure and productivity after additions of naturally occurring dissolved organic matter: results from large-scale lacustrine mesocosms. Limnology and Oceanography. 60, 2130-2144. Link

Fey SB=, Siepielski AM=, Nusslé S, Cervantes-Yoshida K, Hwan JL, Huber ER, Fey MJ*, A Catenazzi, Carlson SM. 2015. Recent shifts in the occurrence, causes, and magnitude of animal mass mortality events. Proceedings of the National Academy of Sciences, USA. 112, 1083-1088. Link    Media Coverage: National GeographicKPCC Public RadioHartford Courant, BloombergCBS,  Hippo ReadsFrontiers in Ecology and the EnvironmentAudubon MagazineThe New Yorker

Fey SB, Mertens AN*, Cottingham KL. 2015. Autumn leaf subsidies influence spring dynamics of freshwater plankton communities. Oecologia. 178, 875-885. Link

Fey SB, Mertens AN*, Beversdorf LJ, McMahon KD, Cottingham KL. 2015. Climate change consequences cross ecosystem boundaries: warming impacts lake plankton communities by modifying terrestrial vegetation subsidies. Oikos. 124, 1473-1481. Link *Cover Image*


Fey SB, Herren CM*. 2014. Temperature-mediated biotic interactions influence enemy release of non-native species in warming environments. Ecology. 95, 2246-2256. Link


Bertilsson S, Burgin A, Carey CC, Fey SB, Grossart H, Grubisic L, Jones I, Kirillin G, Lennon JT, Shade A, Robyn S. 2013. The below-ice microbiome of seasonally frozen lakes.  Limnology and Oceanography. 58, 1998-2012. Link

O’Donnell D*, Fey SB, Cottingham KL. 2013. Nutrient availability influences kairomone-induced defenses in Scenedesmus acutus (Chlorophyceae). Journal of Plankton Research. 35, 191-200. Link

2012 & Earlier

Fey SB, Cottingham KL. 2012. Thermal sensitivity predicts the establishment success of non-native species in a mesocosm warming experiment. Ecology. 93, 2313-2320. Link

Fey SB, Cottingham KL. 2011. Linking biotic interactions and climate change to the success of exotic Daphnia lumholtzi. Freshwater Biology. 56, 2196-2209. Link

Fey SB, Mayer ZA*, Davis SD*, Cottingham KL. 2010. Zooplankton grazing of Gloeotrichia echinulata and associated life history consequences. Journal of Plankton Research. 32, 1337-1347. Link  **2010 David Cushing Prize winner**




Samuel Fey
Biology Department
Reed College
3203 SE Woodstock Blvd.
Portland, OR 97202-8199


Phone: 860-573-8838

CV (Feb 2024, as pdf)

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