1. Allen, W. L., et al. (2011). "Why the leopard got its spots: relating pattern development to ecology in felids." Proceedings of the Royal Society B-Biological Sciences 278(1710): 1373-1380.  

  2. Collier, G. E. and S. J. Obrien (1985). "A Molecular Phylogeny of the Felidae - Immunological Distance." Evolution 39(3): 473-487.  

  3. Day, L. M. and B. C. Jayne (2007). "Interspecific scaling of the morphology and posture of the limbs during the locomotion of cats (Felidae)." Journal of Experimental Biology 210(4): 642-654.  

  4. Hubbard, C., et al. (2009). "Comparative Analysis of Paw Pad Structure in the Clouded Leopard (Neofelis nebulosa) and Domestic Cat (Felis catus)." Anatomical Record-Advances in Integrative Anatomy and Evolutionary Biology 292(8): 1213-1228. Hubbard 2009)

  5. King, L. M. and S. C. Wallace (2014). "Phylogenetics of Panthera, including Panthera atrox, based on craniodental characters." Historical Biology 26(6): 827-833.

  6. Shattuck, M. R. and S. A. Williams (2010). "Arboreality has allowed for the evolution of increased longevity in mammals." Proceedings of the National Academy of Sciences of the United States of America 107(10): 4635-4639.

  7. Sul, S.-J. and T. L. Williams (2011). "Big Cat Phylogenies, Consensus Trees, and Computational Thinking." Journal of Computational Biology 18(7): 895-906.  

  8. Van Valkenburgh, B. (1987). "Skeletal indicators of locomotor behaviour in living and extinct carnivores." Journal of Vertebrate Paleontology 7(2): 162-182.

  9. Vanvalkenburgh, B. (1985). "Locomotor Diversity within Past and Present Guilds of Large Predatory Mammals." Paleobiology 11(4): 406-428.  

  10. Wilting, A., et al. (2007). "Two modern species of clouded leopard: a molecular perspective." Cat News 47: 10-11.  

  11. Martin, P. & Bateson, P. 1985. The influence of experimentally manipulating a component of weaning on the development of play in domestic cats.Anita.Behav.,33, 511-518      

  12. Martin, L. 1980. Functional morphology and the evolution of cats. Transactions of the Nebraska Academy of Sciences, VIII:141-154

  13. Yu, L. Zhang, Y.P. Phylogenetic studies of Pantherine cats (Felidae) based on multiple gene, with novel applications of nuclear beta-fibrinogen intron 7 to carnivores. Mol. Phy. Evol. 2005, 35:483-495

  14. Lynam, A., et al. (2013). Terrestrial activity patterns of wild cats from camera trapping.  THE RAFFLES BULLETIN OF ZOOLOGY 2013 61(1): 407–415

  15. Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)   

  16. Sunquist, F., Sunquist M. Wild Cats of the World. University of Chicago Press. 2002

  17. Bekoff, Marc, and John A. Byers. "A critical reanalysis of the ontogeny and phylogeny of mammalian social and locomotor play: An ethological hornet’s nest." Behavioral development: The Bielefeld interdisciplinary project (1981): 296-337.

  18. Martin, P.  Bateson, P. 1985. The ontogeny of locomotor behavior in the domestic cat. Animal Behavior. Vol. 33. Issue 2, pg 502-510

  19. Clouded Leopard Project, 2003-2011. <>