How does it develop?

In haplodiploid species, an individual's genome and sex determines the role it will play in the population. There is generally one female who reproduces, commonly called the "queen" or "foundress", who contributes DNA to all offspring, male or female. A population under this condition is monogynous.

In one experiment with Augochlorella aurata(Mueller 1994, "Haplodiploidy and the evolution of facultative sex-ratios in a primmitively eusocial bee.") many such colonies were identified and designated "eusocial" or "parasocial". The eusocial colonies were left as is, while the parasocial colonies had the queen removed to be replaced by a single worker. In some cases, a third group was created, where during the course of the experiment a few eusocial colonies underwent natural supersession as the queen died and was replaced.

When the queen was replaced by a new queen that was previously a worker, whether by removal of the queen as part of the experiment or natural death, the sex ratio of the population tended to skew male. This is because of both the new queen's inability to produce diploid offspring without mating as well as her inability to prevent other worker bees from reproducing, which the foundress queen was previously able to do.